Cooke, G. M*. and Turner G. F. Team colours matter when playing away from home: Allopatric aggression in African cichlids. Submitted Hydrobiologica June 2017

Almansa, A.V. Sykes, G. Ponte , G. M. Cooke, P.L.R. Andrews. The digestive tract and welfare of cephalopods in the laboratory and aquaculture. Accepted June 2017  Frontiers in Physiology

Ponte, G., Sykes, A. V., Cooke, G. M., Almansa, E., & Andrews, P. (2017). The digestive tract of cephalopods: towards non-invasive in vivo monitoring of its physiology. Frontiers in Physiology, 8, 403.

Book chapter: Cooke, G.M*, Crackenel, D. and Hill, G.  Public Aquaria Research Guide in BIAZA research methods. Ed. Dr Amy Ploughman, Dr Geoff Hosey

Nicholas R. Casewell, Jeroen Visser, Kate Baumann, James Dobson, Han Han, Michael Morgan, Jordan Debono, Ivan Koludarov, Gregory Bird, Gavan M. Cooke, Amanda Nouwens, Wayne C. Hodgson, Simon C. Wagstaff, Karen Cheney, Irina Vetter, Michael K. Richardson, Sanjaya Kuruppu & Bryan G. Fry. Fish, fangs and foolery: evolution of venom and mimicry systems in coral reef fish.  Current Biology February 2017

Greenway, E. Jones, K. Cooke, G.M* 2016 Environmental Enrichment in Captive Juvenile Thornback Rays, Raja clavata (Linnaeus 1758): Applied Animal Behaviour Science.  182 (86–93)

Cooke, G.M* 2015. Letter to the Editor. Concern regarding use of the term “nonhuman animals” in article titled Ontogeny and phylogeny of facial expression of pain. PAIN. 156(9):1826–1829.

Cooke, G. M* & Tonkins, B. M. 2015 Behavioural indicators of welfare exhibited by the common European cuttlefish (Sepia officinalis). Journal of Zoo and Aquarium Research 3(4)

Tonkins, Belinda M., Alexandra M. Tyers, and Gavan M. Cooke*. 2015 “Cuttlefish in captivity: An investigation into housing and husbandry for improving welfare.” Applied Animal Behaviour Science 168 (77-83).

Gough, R, Hollima, J. Cooke, G.M, Freeman, C. 2015 Characterisation of algogenic organic matter during an algal bloom and its implications for trihalomethane formation. Sustainability of Water Quality and Ecology. 01/2015;

Tyers, A.M., D. Bavin, Cooke, G.M, C. Griggs, and G.F. Turner.  2014.  Peripheral isolate speciation of a Lake Malawi cichlid fish from shallow muddy habitats. Evolutionary Biology doi:10.1007/s11692-014-9277-4

*as corresponding author

In Preparation

Book chapter: Cooke, G.M* and Tonkins, B. M. “Environmental enrichment, cognition and sentience in cephalopod molluscs” in “The Welfare of Invertebrate Animals.”. Ed. Dr Jenifer Mather and Dr Claudio Carere.

Cooke, G. M* Dittrich, A.  Female aggression in African cichlids. Manuscript in preparation for Behavioural Ecology and Sociobiology

Cooke, G.M*, Tonkins, B.M., Ponte, G., Andrews P. L. R. Severity assessment in cephalopod procedures: a consensus approach. Manuscript in preparation for PLOS1

Tyers, A.M, Jones, K. Cooke, G.M*, Reactors and Thinkers: Individual variation in behavioural plasticity crosses contexts in cuttlefish. Manuscript in preparation for Animal Behaviour

*as corresponding author




Slater. M Rundle. R, 2014. Management Guidelines for the Welfare of Zoo and Aquarium Animals; Cuttlefish (Sepia officinalis) BIAZA Aquarium Working Group Cephalopod Focus Group. Edited by Tyson P, and Coulson J; Reviewed by Cooke, G.M

Tyers A, Bell A and Cooke G M. 2012 Investigation into the cause of, and effective treatment for, ‘wasting disease’ in cichlid fish stocks. Bangor University

Turner, G.F & Cooke, G.M 2011. The Welfare Benefits of Physical Isolation of Cichlid Fish. Bangor University

Cooke, G, M. 2012. Bangor University School of Biological Sciences Research Aquarium Management Guidelines 4th ed. p1- 79


  • 6th UK and Ireland SHAPE Environmental Enrichment Conference (REEC) Dublin, May 2016. “Aquatic Animal Enrichment”
  • University of Basel, Switzerland, Evolutionary Biology Seminar Series, April 2010. “The behavioural ecology of aggression in Lake Malawi haplochromine cichlid fish”


  • Aquaculture Europe, 2016, Edinburgh, oral presentation: “Effects of increased volume and tank bottom areas on cuttlefish Sepia officinalis, reproduction behaviour”
  • RSPCA/APHA ‘Welfare of wild animals used in research’ meeting, 2016, oral presentation: “Recent Inclusion of the Class Cephalooda (Mollusca) in ASPA1986/EU Directive 2010/63/EU and The Use of Wild Caught Specimens in Scientific Procedures”
  • LASA/UFAW meeting, 2016, oral presentation: Welfare assessment in alien life forms: The challenges presented by inclusion of cephalopods in directive 2010/63/EU
  • BBSRC Animal Welfare Research Network Inaugural Meeting, May 2016, Welcome Trust, London, poster: “COST Action FA1301 – CephsInAction A network for improvement of cephalopod welfare and husbandry in research, aquaculture and fisheries”
  • FELASA congress 2016, Brussels, poster: “Prospective severity assessment in cephalopods: Results of an on-line survey of the COST Action (CephsInAction) FA1301 cephalopod research community”
  • Aquaculture Europe, 2015, Rotterdam, oral presentation: “Caring for cephalopods”
  • BIAZA Research Conference 2015, Dublin, oral presentation: “Improving the lives of captive Rays”
  • NAW BIAZA Conference, 2014, Liverpool, oral presentation: “The science of animal welfare: caring for cuttlefish”
  • 9th World Congress on Alternatives and Animal Use in the Life Sciences 2014, Prague, poster, “Husbandry and Enrichment for the Recently Protected Common European Cuttlefish”
  • Cichlid Science 2013 Bangor – conference organiser and oral presentation: “Husbandry Techniques in Cichlid Research”
  • Cichlid Science 2012 Leuven, Belgium, oral presentation: “Aggression in Malawi Mbuna cichlids”
  • Cichlid Science 2010 Basel, Switzerland, oral presentation: “Can Satellite lakes act as species nurseries?”



  • UFAW Summer studentship award (as supervisor, captive Thornback Ray enrichment), 2015 £1600
  • Universities Federation for Animal Welfare (UFAW) grant, 2014, £800 to present welfare research at 9th World Congress for Animal Welfare (Prague)
  • Bangor University internship scheme (as supervisor, Cuttlefish welfare and enrichments), 2014 £1500
  • ZSL Erasmus Darwin Grant 2009, £4000 conservation field work in Africa


PADI open water certificate

Powerboat level 2 certificate

First Aid in the field certificate

2 X 6 month deployment living aboard HM RN survey ship HMS Beagle as hydrographic surveyor technician (Plymouth to Arabian Gulf)

3 x 28 day field work in an around Lake Malawi using using underwater behvaioural observations, lake limnology techniques and supervision of groups of students

Vast experience of UK coastal,  estuarine,  lake and riverine habitats and familiar with a wide variety of survey aquatic (biotic/abiotic) techniques