Academic Curriculum Vitae

Dr. Gavan M Cooke BSc (Hons.) PhD PG Cert. Ed


  • PGCert in Education Anglia Ruskin University, UK – 2016-2017
  • Doctorate (Behavioural Ecology/Evolutionary Biology of cichlid fish) Bangor University, Bangor, Wales, UK 2007-2012
  • Bachelor’s Degree – Zoology BSc (Hons)University of Leeds, UK 2002-2006


The Cephalopod Citizen Science Project – Dec 2017 – present

Creator, owner and director of a global (14 countries, 3000 members) cephalopod citizen science project – please see here for the scope and overview of the project The Cephalopod Citizen Science Project 

Anglia Ruskin University

Lecturer in Animal Behaviour Sept 2016 – present

  • Teaching at an undergraduate and postgraduate level using ‘team-based learning’, readiness assessment tests and classical higher education teaching methods (lecturing, practical sessions and field trips)
  • Module manager for level 6 modules: Human-Animal Management for Improving Welfare; Animal Heath and Nutrition;  Animal Learning and Training.
  • Lectured on other modules such as Practical Skills for Animal Behaviourists and Management of Zoo Animals. Undergraduate and Masters project supervision
  • Chair of the ethical review committee

The Cornwall College Group (Falmouth Marine School)

Course leader Aquaculture and Marine Science Jul 2015 to Mar 2016

Bangor University, Bangor, Wales, UK

Aquarium Facility Manager/lecturer/researcher Jun 2010 to Jul 2015

  • Significant role in design and construction of 125k litre marine and freshwater research aquarium.
  • Responsible for all aspects of the animal facility including animal welfare, H&S, staff training etc.
  • Held over 50 freshwater fish species, 4 amphibian species, 15 marine fish species, and 2 cephalopod species and many more marine invertebrates (inc. corals).
  • Winner of the 2014 Bangor University Student Led Teaching Award.

Bangor University, Bangor, Wales,

Demonstrator/lecturer Sept 2007 to Sept 2010

University of Hull, Hull.

Animal Technician Feb 2007 to Sept 2007

Institute for Estuarine and Coastal Studies, Hull

Benthic Technician Sept 2006 to Feb 2007   

Hydrographic Surveyor Technician

HM Royal Navy Jun 1998 to Jun 2000

  • Maintenance of marine science equipment (e.g. echo sounders, side-scan sonar, CDT probes etc.)
  • Entailed detailed record-keeping, data entry, data analysis and the creation of Admiralty charts
  • Developed an ability to work carefully and accurately under extreme
  • Leadership training
  • Learned detailed aspects of sailing and general seamanship.


* As corresponding author

Tyers, Alexandra M., Gavan M. Cooke, and George F. Turner. “Rare morph Lake Malawi mbuna cichlids benefit from reduced aggression from con-and hetero-specifics.” bioRxiv (2021).

Gordon, D., Pugh, P. and Cooke, G.M., 2019*. Social Media and citizen science provide valuable data for behavioural ecology research: Are cuttlefish using pursuit-deterrent signals during hunting?. bioRxiv, p.760926.

Oliveira, A.R., Sardinha-Silva, A., Andrews, P.L., Green, D., Cooke, G.M., Hall, S., Blackburn, K. and Sykes, A.V., 2020. Microplastics presence in cultured and wild-caught cuttlefish, Sepia officinalisMarine Pollution Bulletin160, p.111553.

Sykes, A.V., Almansa, E., Ponte, G., Cooke, G.M. and Andrews, P.L., 2020. Can cephalopods vomit? Hypothesis based on a review of circumstantial evidence and preliminary experimental observations. Frontiers in Physiology11.

Drerup, C., Sykes, A.V. and Cooke, G.M., 2020. Behavioural aspects of the spotty bobtail squid Euprymna parva (Cephalopoda: Sepiolidae). Journal of Experimental Marine Biology and Ecology530, p.151442.

Cooke, G.M*., Anderson, D.B., Begout, M.L., Dennison, N., Osorio, D., Tonkins, B., Kristiansen, T., Fiorito, G., Galligioni, V., Ponte, G. and Andrews, P.L., 2019. Prospective severity classification of scientific procedures in cephalopods: Report of a COST FA1301 Working Group survey. Laboratory animals, p.0023677219864626.

Marshall, L., McCormick, W.D. and Cooke, G.M., 2019. Perception of the ethical acceptability of live prey feeding to aquatic species kept in captivity. PloS one14(8), p.e0216777.

Cooke, G.M*., Tonkins, B.M. and Mather, J.A., 2019. Care and Enrichment for Captive Cephalopods. In The Welfare of Invertebrate Animals (pp. 179-208). Springer, Cham.

Cooke, G.M* and Turner, G.F., 2018. Team colours matter when playing away from home: aggression biases in geographically isolated mbuna cichlid populations. Hydrobiologia, 809(1), pp.31-40.

Cooke, G.M.*, 2017. Stereotypic behaviour is not limited to terrestrial taxa: A response to Rose et al. Journal of Veterinary Behavior: Clinical Applications and Research, 22, pp.17-18.

Cooke, G.M.*, 2017. Use of a translucent refuge for Xenopus tropicalis with the aim of improving welfare. Laboratory animals, p.0023677217737798.

Sykes, António V., Eduardo Almansa, Gavan M. Cooke, Giovanna Ponte, and Paul LR Andrews. “The digestive tract of cephalopods: a neglected topic of relevance to animal welfare in the laboratory and aquaculture.” Frontiers in Physiology 8 (2017): 492.

Ponte, G., Sykes, A. V., Cooke, G. M., Almansa, E., & Andrews, P. (2017). The digestive tract of cephalopods: towards non-invasive in vivo monitoring of its physiology. Frontiers in Physiology, 8, 403.

Book chapter: Cooke, G.M*, Cracknell, D. and Hill, G.  (2017) Public Aquaria Research Guide in BIAZA research methods. Ed. Dr Amy Ploughman, Dr Geoff Hosey

Nicholas R. Casewell, Jeroen Visser, Kate Baumann, James Dobson, Han Han, Michael Morgan, Jordan Debono, Ivan Koludarov, Gregory Bird, Gavan M. Cooke, Amanda Nouwens, Wayne C. Hodgson, Simon C. Wagstaff, Karen Cheney, Irina Vetter, Michael K. Richardson, Sanjaya Kuruppu & Bryan G. Fry. Fish, fangs and foolery: evolution of venom and mimicry systems in coral reef fish.  Current Biology February 2017

Greenway, E. Jones, K. Cooke, G.M* 2016 Environmental Enrichment in Captive Juvenile Thornback Rays, Raja clavata (Linnaeus 1758): Applied Animal Behaviour Science.  182 (86–93)

Cooke, G.M* 2015. Letter to the Editor. Concern regarding use of the term “nonhuman animals” in the article titled Ontogeny and phylogeny of facial expression of pain. PAIN. 156(9):1826–1829.

Cooke, G. M* & Tonkins, B. M. 2015 Behavioural indicators of welfare exhibited by the common European cuttlefish (Sepia officinalis). Journal of Zoo and Aquarium Research 3(4)

Tonkins, Belinda M., Alexandra M. Tyers, and Gavan M. Cooke*. 2015 “Cuttlefish in captivity: An investigation into housing and husbandry for improving welfare.” Applied Animal Behaviour Science 168 (77-83).

Gough, R, Holliman, J. Cooke, G.M, Freeman, C. 2015 Characterisation of allogenic organic matter during an algal bloom and its implications for trihalomethane formation. Sustainability of Water Quality and Ecology. 01/2015;

Tyers, A.M., D. Bavin, Cooke, G.M, C. Griggs, and G.F. Turner.  2014.  Peripheral isolate speciation of a Lake Malawi cichlid fish from shallow muddy habitats. Evolutionary Biology


  • 6th UK and Ireland SHAPE Environmental Enrichment Conference (REEC) Dublin, May 2016. “Aquatic Animal Enrichment”
  • University of Basel, Switzerland, Evolutionary Biology Seminar Series, April 2010. “The behavioural ecology of aggression in Lake Malawi haplochromine cichlid fish”


  • CIAC/Cephs in Action conference, Heraklion, Crete March 2017 Presenting talk: “Prospective severity assessment in cephalopods: Results of an on-line survey of the COST Action (CephsInAction) FA1301 cephalopod research community”
  • Aquaculture Europe, 2016, Edinburgh, oral presentation: “Effects of increased volume and tank bottom areas on cuttlefish Sepia officinalis, reproduction behaviour”
  • RSPCA/APHA ‘Welfare of wild animals used in research’ meeting, 2016, oral presentation: “Recent Inclusion of the Class Cephalopoda (Mollusca) in ASPA1986/EU Directive 2010/63/EU and The Use of Wild Caught Specimens in Scientific Procedures”
  • LASA/UFAW meeting, 2016, oral presentation: Welfare assessment in alien life forms: The challenges presented by inclusion of cephalopods in directive 2010/63/EU
  • BBSRC Animal Welfare Research Network Inaugural Meeting, May 2016, Welcome Trust, London, poster: “COST Action FA1301 – CephsInAction A network for improvement of cephalopod welfare and husbandry in research, aquaculture and fisheries”
  • FELASA congress 2016, Brussels, poster: “Prospective severity assessment in cephalopods: Results of an on-line survey of the COST Action (CephsInAction) FA1301 cephalopod research community”
  • Aquaculture Europe, 2015, Rotterdam, oral presentation: “Caring for cephalopods”
  • BIAZA Research Conference 2015, Dublin, oral presentation: “Improving the lives of captive Rays”
  • NAW BIAZA Conference, 2014, Liverpool, oral presentation: “The science of animal welfare: caring for cuttlefish”
  • 9th World Congress on Alternatives and Animal Use in the Life Sciences 2014, Prague, poster, “Husbandry and Enrichment for the Recently Protected Common European Cuttlefish”
  • Cichlid Science 2013 Bangor – conference organiser and oral presentation: “Husbandry Techniques in Cichlid Research”
  • Cichlid Science 2012 Leuven, Belgium, oral presentation: “Aggression in Malawi Mbuna cichlids”
  • Cichlid Science 2010 Basel, Switzerland, oral presentation: “Can Satellite lakes act as species nurseries?”



  • National Geographic SEE Initiative ROV Drone grant win. Value ~$1600 The secret Lives of shell dwelling octopuses
  • National Geographic SEE Initiative ROV Drone grant win. Value ~$1600 How to film cephalopods
  • Anglia Ruskin University undergraduate research taster (as supervisor, Aquaculture welfare), 2017 £2000
  • UFAW Summer studentship award (as supervisor, captive Thornback Ray enrichment), 2015 £1600
  • Universities Federation for Animal Welfare (UFAW) grant, 2014, £800 to present welfare research at 9th World Congress for Animal Welfare (Prague)
  • Bangor University internship scheme (as supervisor, Cuttlefish welfare and enrichments), 2014 £1500
  • ZSL Erasmus Darwin Grant 2009, £4000 conservation fieldwork in Africa




  • Powerboat level 2           August 2015
  • PADI Open Water           February 2015
  • Marine Mammal Medic               March 2014
  • Institute for Animal Technology Level 2 February 2014
  • First Aid in Work             June 2013
UK Home Office (ASPA, 1986) Accredited (Society of Biology) Animal Welfare Training
Named Animal Welfare Officer March 2013
Fish modules 1 (law), 2 (husbandry), 3 (anaesthesia) and 4 (minor surgery) September 2012
Cephalopod modules 1 (law) and 2 (husbandry) August 2012
Bird modules 1, 2, 3 (law, husbandry and anaesthesia) July 2011
Mammal modules, 1, 2, 3 (law, husbandry, and anaesthesia) June 2010